Cephalopods and their ink-producing glands. These 19th century images have been relabeled and retouched for clarity. (A,B) Squid and its ink sac. (A) Illex illecebrosus, from of Plate XXXIX of Verrill in 1880 []. (B) Doryteuthis pealeii, from Figure 13 of Williams in 1909 []. The ink sac is labeled by an asterisk; (C) Ink sac from the cuttlefish, Sepia officinalis, from Figure 291 of Lang and Hescheler in 1900 []. a, anus; 1, common sac of the rectum and ink sac; 2, rectum; 3, ampulla of the duct of the ink sac; 4, anterior sphincter muscle of the ink sac; 5, posterior sphincter muscle of the ink sac; 6, duct of the ink sac; 7, reservoir of the ink sac; 8, opening of the ink gland; 9, ink gland; 10, basal germinal zone; (D,E) Funnel organ of squid; (D) Ventral view of the funnel organ from Desmoteuthis tenera (=Teuthowenia megalops) from d of Plate LV from Verrill in 1880 [] and from Loligo; (E-1) Ventral view and (E-2) dorsal view, from of Laurie in 1888 []. The funnel organ, located on the funnel near the siphon (s) and siphonal valve (v), is composed of a median organ (m), two lateral cushions (n, n′) and a papilla to each (one medio-dorsal papilla, i, and two lateral papillae, i′).