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SRX765371: GSM1551914: Adelman_Bl6_mESC_C2cells_2i_Ctrl_startRNA-seq_rep3; Mus musculus; RNA-Seq
3 ILLUMINA (Illumina MiSeq) runs: 61M spots, 6.2G bases, 2.5Gb downloads

Submitted by: Gene Expression Omnibus (GEO)
Study: Pausing of RNA polymerase II regulates mammalian developmental potential
show Abstracthide Abstract
The remarkable capacity for pluripotency and self-renewal in embryonic stem cells (ESCs) requires a finely-tuned transcriptional circuitry wherein the genes and pathways that initiate differentiation are suppressed, but poised to respond rapidly to developmental signals. To elucidate transcriptional control in mouse ESCs in the naïve, ground state, we defined the distribution of engaged RNA polymerase II (Pol II) at high-resolution. We find that promoter-proximal pausing of Pol II is widespread at genes encoding signaling molecules that control ESC proliferation, differentiation and metabolism, but is not enriched at developmental control genes. Accordingly, ablation of the primary pause-inducing factor, NELF, causes proliferation defects, early embryonic lethality and dysregulation of critical ESC signaling pathways. Moreover, loss of NELF in ESCs causes resistance to cell fate commitment through dramatic attenuation of FGF/ERK activity. This work thus uncovers an unanticipated role for NELF-mediated pausing in establishing the responsiveness of ESCs to developmental cues. Overall design: RNA-seq, GRO-seq, and Start-RNA-seq from mouse embryonic stem cells (mESCs). For RNA-seq, two biological replicates are included for each of two treatment conditions; control, and 4-hydroxytamoxifen (4OHT) treated to ablate expression of NELF-B. For GRO-seq, data from two biological replicates of untreated mESCs are included. For Start-RNA-seq, three biological replicates are included for each of two treatment conditions; control and 4OHT.
Sample: Adelman_Bl6_mESC_C2cells_2i_Ctrl_startRNA-seq_rep3
SAMN03216558 • SRS749670 • All experiments • All runs
Organism: Mus musculus
Library:
Instrument: Illumina MiSeq
Strategy: RNA-Seq
Source: TRANSCRIPTOMIC
Selection: cDNA
Layout: PAIRED
Construction protocol: GRO-seq: GRO-seq experiments were performed according to {Core, 2008 #1330} in the C2 ESC line grown in 2i media, with the following modifications: Nuclei were harvested by swelling trypsinized cells for 10 minutes in 50 mL cold lysis buffer (10 mM Tris pH 7.5, 10 mM NaCl, 3 mM CaCl2, 2 mM MgCl2, 0.5% NP40, 5 mM DTT, 300 mM sucrose, 1 mM PMSF, 2 U/ml Superase IN, 5 tablets Roche protease inhibitors), then gently dounce homogenized, centrifuged and washed once with 50 mL lysis buffer. Nuclei were resuspended in freezing buffer as in {Core, 2008 #1330} at the concentration of 5x106 nuclei per 100 uL, and stored at -80°C until use. To avoid a bias against representation of C-rich regions in the data set, the final concentration of CTP (Roche) was increased in the run-on ten-fold above that used in {Core, 2008 #1330} to 10 uM. To permit a quick, reproducible stop to the run-on reaction, the reaction was terminated after 5 minutes by the addition of Trizol (Invitrogen). RNA-seq: Two distinct conditional NELF-B KO ESC clones were grown +/- 4OHT in 2i media. On day 5 after initiating recombination, total RNA was prepared using Trizol (Invitrogen) and manufacturer recommended methods. DNA contamination was removed using the RNeasy Mini Kit (Qiagen) per manufacturer’s instructions, and RNA quality was assessed using a Bioanalyser Nano ChIP (Agilent). startRNA-seq: Total RNA was extracted from nuclei using Trizol reagent (Invitrogen) GRO-seq: Following base hydrolysis, primary Br-U immunoprecipitation and end repair, the libraries were made using the Illumina small RNA TruSeq kit except the 3’ adapter was ligated using T4 RNA Ligase II, truncated KQ enzyme (NEB). A secondary BrU IP was performed before ligation of the 5’ adapter (Illumina) followed by a tertiary BrU IP to produce triple-selected run-on RNA. RNA was reverse transcribed and amplified with 15 cycles of PCR (Illumina). Libraries were PAGE purified, selecting for products larger than 140 bp. RNA-seq: Ribosomal RNA was removed prior to library construction by hybridizing to ribo-depletion beads that contain biotinylated capture probes (Ribo-Zero, Epicentre). RNA was then fragmented and libraries were prepared according to the TruSeq Stranded Total RNA Gold Kit (Illumina) using random hexamer priming. Illumina Spike-ins were used for normalization. startRNA-seq: Start-RNA libraries were prepared as described in (Nechaev et al., Science 2010) except reads were size selected in the range 20-80 nt to exclude full length snRNA species.
Experiment attributes:
GEO Accession: GSM1551914
Links:
External link:
Runs: 3 runs, 61M spots, 6.2G bases, 2.5Gb
Run# of Spots# of BasesSizePublished
SRR1973374244,18624.4M11.9Mb2015-04-14
SRR19733758,676,196867.6M389.3Mb2015-04-14
SRR197337652,061,2625.3G2.1Gb2015-04-14

ID:
1105294

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