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SRX5179251: GSM3530162: E14-serum_2; Mus musculus; Hi-C
2 ILLUMINA (Illumina HiSeq 4000) runs: 583.1M spots, 115.4G bases, 41.5Gb downloads

Submitted by: NCBI (GEO)
Study: Hi-C of WT E14 mouse ES cells grown in serum and 2i media
show Abstracthide Abstract
Dynamic reprogramming of global DNA methylation impacts on the genomic deposition of the Polycomb-mediated repressive histone mark H3K27me3. DNA hypomethylation in ground state embryonic stem cells (ESCs) results in a reversible redistribution of H3K27me3 from its normal target loci. Thus, a signalling induced shift of ESCs to ground state results in both DNA methylation and Polycomb patterns that are quite distinct from their primed counterparts. Here we investigated the impact of DNA methylation directed Polycomb redistribution on higher-order chromatin structure in the ground state. Using a targeted single-locus approach (FISH) we can demonstrate local decompaction at Hox loci in the ground state, which is consistent with genome-wide data (Hi-C) indicating that chromatin structure is globally altered in ground state relative to primed ESCs. Polycomb targets are similarly decompacted in hypomethylated E3.5 mouse blastocysts. ESC lines which maintain a high level of DNA methylation in ground state show no decompaction at Hox loci. Our results suggest that DNA-methylation mediated reprogramming of Polycomb binding drives higher order chromatin organisation in stem cells and early development. Overall design: In situ Hi-C experiments on E14 mESCs in serum and 2i culture with two replicates per condition
Sample: E14-serum_2
SAMN10642588 • SRS4186504 • All experiments • All runs
Organism: Mus musculus
Library:
Instrument: Illumina HiSeq 4000
Strategy: Hi-C
Source: GENOMIC
Selection: other
Layout: PAIRED
Construction protocol: Hi-C was performed largely as described (Rao et. el., 2014) with minor modifications. Briefly, ES cells were crosslinked in 1% formaldehyde for 10 minutes, snap-frozen in 2-5 mln aliquots and stored at -80°C. After permeabilization in lysis buffer (0.2% Igepal, 10 mM Tris-HCl pH 8.0, 10 mM NaCl, 1x Halt Protease inhibitor cocktail) nuclei were isolated in 0.3% SDS in NEBuffer 3 at 62° for 10 minutes. SDS was quenched with 1% Triton X-100 at 37° for 1 h, then the nuclei were pelleted and resuspended in 250 ul DpnII buffer with 600 U DpnII. After overnight digestion, 200 more units were added for 2 hours. Then the ends were filled-in using Klenow, d(G/C/T)TPs and biotin-14-dATP for 1.5 hrs at 37°. After ligation at room temperature for 4 hrs the nuclei were spun down, resuspended in 200 ul mQ and digested with proteinase K for 30 min at 55° in presence of 1% SDS. Then the cross-links were reversed at 65° overnight after addition of NaCl to a final concentration of 1.85 M. In the morning, after ethanol precipitation and a wash with 70-80% Ethanol, DNA was resuspended in 500 ul of sonication buffer (50 mM Tris pH 8.0, 0.1%SDS, 10 mM EDTA), incubated on ice for 15 min and then sheared using a probe sonicator to fragment size of 200-700 bp. Then the DNA was concentrated on Amicon filter units, bound to MyOne T1 Streptavidin beads and used for Illumina library preparation. Small aliquots were taken before and after DpnII treatment, and before sonication to confirm efficient DNA digestion and ligation by running them on 1% agarose gel. Samples were first test-sequenced on NextSeq 550 (WTCRF, Edinburgh) to check library quality, and then selected libraries were sequenced deeply on HiSeq 4000 (BGI). In house Illumina library protocol from Rao et. al., 2014.
Experiment attributes:
GEO Accession: GSM3530162
Links:
Runs: 2 runs, 583.1M spots, 115.4G bases, 41.5Gb
Run# of Spots# of BasesSizePublished
SRR836906325,447,7613.8G1.7Gb2019-02-04
SRR8369064557,605,225111.5G39.8Gb2019-02-04

ID:
6994278

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