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SRX2708709: GSM2563787: nasBS-seq_pulse_parent_rep6; Homo sapiens; Bisulfite-Seq
1 ILLUMINA (Illumina HiSeq 2500) run: 259.5M spots, 25.9G bases, 8.9Gb downloads

Submitted by: NCBI (GEO)
Study: Mapping DNA methylation and CTCF/cohesin occupancy on nascent chromatin and DNMT-targeted nascent chromatin
show Abstracthide Abstract
The faithful inheritance of the epigenome is critical for cells to maintain gene expression programs and cellular identity across cell divisions. We mapped strand-specific DNA methylation after replication forks and show maintenance of the vast majority of the DNA methylome within 20 minutes of replication and inheritance of some hemimethylated CpG dinucleotides (hemiCpGs). Mapping the nascent DNA methylome targeted by each of the three DNA methyltransferases (DNMTs) reveals interactions between DNMTs and substrate daughter cytosines en route to maintenance methylation or hemimethylation. Finally, we show the inheritance of hemiCpGs at short regions flanking CCCTC-binding factor (CTCF)/cohesin binding sites in pluripotent cells. Elimination of hemimethylation causes reduced frequency of chromatin interactions emanating from these sites, suggesting a role for hemimethylation as a stable epigenetic mark regulating CTCF-mediated chromatin interactions. Overall design: Mapping DNA methylation and CTCF/cohesin occupancy on nascent chromatin and DNMT-targeted nascent chromatin
Sample: nasBS-seq_pulse_parent_rep6
SAMN06685691 • SRS2101416 • All experiments • All runs
Organism: Homo sapiens
Library:
Instrument: Illumina HiSeq 2500
Strategy: Bisulfite-Seq
Source: GENOMIC
Selection: RANDOM
Layout: PAIRED
Construction protocol: For pulse, H9 cells were transferred to medium containing 50 µM EdU and were cultured at 37 °C for 20 min before formaldehyde crosslinking. For chase, after labeling with EdU, cells were washed with DPBS twice, and cultured in medium containing 50 µM dT for 8 hr before formaldehyde crosslinking. Crosslinking with 1% formaldehyde was done in DPBS at RT for 10 min and was quenched with 0.125 M glycine. Cells were collected and washed with DPBS, and lysed in Lysis Buffer (5 mM PIPES pH 8.0, 85 mM KCl, 0.5% NP-40, 10% glycerol). Pellets were collected and re-suspended in Sonication Buffer (10 mM Tris-HCl pH 8.0, 100 mM NaCl, 1 mM EDTA pH 8.0, 0.5 mM EGTA pH 8.0, 0.1% sodium deoxycholate, 0.5% sodium lauroylsarcosine). Sonication was done on ice using Bioruptor. Triton X-100 was added to a final concentration of 1% before centrifugation at 15,000 xg. Supernatant was collected as fraction of chromatin. Elution Buffer (50 mM Tris-HCl pH 8.0, 10 mM EDTA pH 8.0, 1% SDS) was added to chromatin in a 1:1 ratio. Crosslinking reversal was done by incubation at 65 °C for 8 hr. Proteinase K was added to a final concentration of 0.2 mg/ml before incubation at 55 °C for 30 min. Phenol:chloroform:isoamyl alcohol extraction and ethanol precipitation was done sequentially to purify genomic DNA. Click chemistry was performed in a 200 µl volume (174 µl DNA in DPBS, 2 µl 1 mM biotin-azide, 4 µl 100 mM CuSO4, 20 µl 100 mM sodium ascorbate) at 37 °C for 1 hr. After ethanol precipitation, DNA was incubated with Dynabeads Streptavidin C1 beads on a rotor for 30 min at RT, and washed according to the manufacturer’s instructions. Beads were incubated with End Repair Enzyme Mix on a rotor for 30 min at RT. After washing, beads were incubated with Klenow Fragment (3'→5' exo-) reaction mix on a rotor for 30 min at RT. After washing, beads were incubated with T4 DNA ligase reaction mix and methylated adaptors on a rotor for 2 hr at RT. After washing, beads were incubated in 150 mM NaOH for 3 min at RT. Supernatant was collected as the fraction containing the parental strand DNA (bio-). Beads were washed 3 times with NaOH, re-suspended in 95% formamide and 10 mM EDTA pH 8.2, and incubated at 90 °C for 3 min. Supernatant was collected as the fraction containing the daughter strand DNA (bio+). Both fractions were ethanol precipitated, mixed with 1 pg of lambda DNA, and bisulfite converted with the EZ DNA Methylation-Lightning Kit. PCR amplification was done with HiFi Uracil+ polymerase. For each library, the minimal number of PCR cycles were used to yield at least 20 ng of DNA measured after purification with AMpure XP beads.
Experiment attributes:
GEO Accession: GSM2563787
Links:
Runs: 1 run, 259.5M spots, 25.9G bases, 8.9Gb
Run# of Spots# of BasesSizePublished
SRR5417115259,492,47025.9G8.9Gb2018-03-13

ID:
3908565

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