1 Department of Cell Biology and Anatomy and the Alberta Children's Hospital Research Institute, University of Calgary, Calgary, Alberta, Canada.
2 Department of Mathematics, Florida State University, Tallahassee, Florida, United States of America.
3 Department of Orthopedics, University of California, San Francisco, San Francisco, California, United States of America.
4 Department of Pediatrics and Human Medical Genetics and Genomics Program, University of Colorado School of Medicine, Denver, Colorado, United States of America.