1 Department of Biology, University of Massachusetts, Amherst, MA, United States. Electronic address: mfacette@umass.edu.
2 Department of Botany and Plant Sciences, Center for Plant Cell Biology, Institute of Integrative Genome Biology, University of California, Riverside, CA, United States. Electronic address: carolyn.rasmussen@ucr.edu.
3 Department of Botany and Plant Sciences, Center for Plant Cell Biology, Institute of Integrative Genome Biology, University of California, Riverside, CA, United States. Electronic address: jaimie.vannorman@ucr.edu.