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Zootaxa. 2014 Oct 1;3869(3):201-23. doi: 10.11646/zootaxa.3869.3.1.

A new toothed pterosaur (Pterodactyloidea: Anhangueridae) from the Early Cretaceous Romualdo Formation, NE Brazil.

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Programa de Pós-graduação em Geociências, Centro de Tecnologia e Geociências, Universidade Federal de Pernambuco, Av. Acadêmico Hélio Ramos s/n, Cidade Universitária, Recife, Pernambuco, Brazil.; Email:
Laboratório de Paleontologia, Universidade Regional do Cariri, Rua Carolino Sucupira s/n, Pimenta, Crato, Ceará, Brazil.; Email:
Laboratório de Paleontologia e Sistemática, Departamento de Biologia, Universidade Federal Rural de Pernambuco, Rua Dom Manuel de Medeiros s/n, Dois Irmãos, Recife, Pernambuco, Brazil.; Email:
Laboratório de Biodiversidade do Nordeste, Universidade Federal de Pernambuco, Rua do Alto Reservatório s/n, Bela Vista, Vitória de Santo Antão, Pernambuco, Brazil.; Email:


A new species of pterosaur, Maaradactylus kellneri gen. nov., sp. nov. (Archosauria: Pterosauria) from the Romualdo Formation (Aptian/Albian), is herein described. The specimen (MPSC R 2357) was found at Sítio São Gonçalo, Santana do Cariri city (State of Ceará, northeast Brazil) and consists of the skull, atlas and axis, and represents one of the largest skulls of the Anhangueridae from the Araripe Basin described. The autapomorphies of the new pterosaur include the following characters: a premaxillary sagittal crest that is relatively long and high, beginning at the anterior part of the skull (rostrum) and extending to the 22nd pair of alveoli, not covering the nasoantorbital fenestra or the choanaes, and also the presence of 35 pairs of alveoli; smooth palatal ridge, which starts on the 5th pair of alveoli and ends on the 13th pair; palate is convex shaped in the anterior region; choanae not extending laterally; small and convex palatal elevation; the 5th, 6th and 7th alveoli smaller than the 4th and 8th; the alveoli decreasing in size from the 9th to the 12th and increasing from the 13th to 18th, and from the 18th to the 35th they are arranged in triplets. Furthermore, the lateral surface of the premaxillary crest shows grooves and tridimensional structures that may have housed blood vessels.

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