PMC

Bayesian tree for the 27 sequences of Lithobiomorpha based on COI sequences. The Bayesian posterior probabilities from Bayesian analyses are presented above the main branches. The scale bar represents substitutions per site.

Distribution of Hessebius Verhoeff, 1941 in China. Symbols denote localities and species. Star = Xinjiang Uygur H longispinipes Ma, Pei and Zhu, 2009; circle = Tibet H multiforaminis Pei, Ma, Zapparoli and Zhu, 2010; triangle = Sichuan H jangtseanus (Verhoeff, 1942); pentagon = Luqu H luquensis sp. n.; square = Sichuan H ruoergaiensis sp. n.

Hessebius luquensis sp. n., holotype, female: A dorsal view, scale 1 mm B left ocelli and Tömösváry’s organ (To), scale 250 μm; C, forcipular coxosternite, ventral view, scale bar 1 mm D posterior segments and gonopods, internal view, scale bar 500 μm E right gonopod, dorsal view, scale bar 250 μm F, right gonopod, ventral lateral view, scale bar 250 μm. Paratype, LQ 9, male G posterior segments and gonopods, internal view, scale bar 500 μm.

Hessebius ruoergaiensis sp. n., holotype, female: A dorsal view, scale bar 1 mm B left ocelli and Tömösváry’s organ (To), scale bar 250 μm D forcipular coxosternite, ventral view, scale bar 1 mm E right gonopod, dorsal view, scale bar 250 μm C, F–H paratype, C, REG1, female: left ocelli and Tömösváry’s organ (To), scale bar 250 μm F REG5, female: right gonopod, ventral lateral view, scale bar 250 μm G REG6, male: male left leg 15, dorsal lateral view, scale bar 500 μm H REG10, male: posterior segments and gonopods, internal view, scale bar 500 μm.