H(+)-transporting ATP synthase, vacuolar type, subunit D
Although this ATPase can run backwards, using a proton gradient to synthesize ATP, the primary biological role is to acidify some compartment, such as yeast vacuole (a lysosomal homolog) or the interior of a prokaryote. [Transport and binding proteins, Cations and iron carrying compounds]
Jiyao W et al.(2023). "The conserved domain database in 2023.",