This domain is found at the N-terminus of P74 occlusion-derived virus (ODV) envelope proteins which are required for oral infectivity. The envelope proteins are found in baculoviruses which are insect pathogens. The C-terminus of P74 is anchored to the membrane whereas the N-terminus is exposed to the virion surface. Furthermore P74 is unusual for a virus envelope protein as it lacks an N-terminal localization signal sequence. Also see pfam04583.
Jiyao W et al.(2023). "The conserved domain database in 2023.",