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Soluble CD137 (4-1BB) ligand is released following leukocyte activation and is found in sera of patients with hematological malignancies.

Salih HR, Schmetzer HM, Burke C, Starling GC, Dunn R, Pelka-Fleischer R, Nuessler V, Kiener PA.

J Immunol. 2001 Oct 1;167(7):4059-66.


Changes in lymphocytes' telomerase activity by 4-1BB costimulation.

Ahmad HT, Mansooreh J, Fereshteh M, Mojtaba H.

J Cancer Res Ther. 2014 Oct-Dec;10(4):998-1003. doi: 10.4103/0973-1482.137906.


4-1BB ligand signaling to T cells limits T cell activation.

Eun SY, Lee SW, Xu Y, Croft M.

J Immunol. 2015 Jan 1;194(1):134-41. doi: 10.4049/jimmunol.1401383. Epub 2014 Nov 17.


Involvement of the 4-1BB/4-1BBL pathway in control of monocyte numbers by invariant NKT cells.

Cole SL, Benam KH, McMichael AJ, Ho LP.

J Immunol. 2014 Apr 15;192(8):3898-907. doi: 10.4049/jimmunol.1302385. Epub 2014 Mar 17.


Blockade of 4-1BB and 4-1BBL interaction reduces obesity-induced skeletal muscle inflammation.

Le NH, Kim CS, Tu TH, Choi HS, Kim BS, Kawada T, Goto T, Park T, Park JH, Yu R.

Mediators Inflamm. 2013;2013:865159. doi: 10.1155/2013/865159. Epub 2013 Dec 18.


Combining antibody-directed presentation of IL-15 and 4-1BBL in a trifunctional fusion protein for cancer immunotherapy.

Kermer V, Hornig N, Harder M, Bondarieva A, Kontermann RE, Müller D.

Mol Cancer Ther. 2014 Jan;13(1):112-21. doi: 10.1158/1535-7163.MCT-13-0282. Epub 2013 Nov 6.


Enhanced antitumor activity mediated by human 4-1BB-engineered T cells.

Daniel-Meshulam I, Horovitz-Fried M, Cohen CJ.

Int J Cancer. 2013 Dec 15;133(12):2903-13. doi: 10.1002/ijc.28320. Epub 2013 Jul 16.


The TNF receptor-ligands 4-1BB-4-1BBL and GITR-GITRL in NK cell responses.

Barao I.

Front Immunol. 2013 Jan 4;3:402. doi: 10.3389/fimmu.2012.00402. eCollection 2012.


4-1BB/4-1BBL interaction promotes obesity-induced adipose inflammation by triggering bidirectional inflammatory signaling in adipocytes/macrophages.

Tu TH, Kim CS, Goto T, Kawada T, Kim BS, Yu R.

Mediators Inflamm. 2012;2012:972629. doi: 10.1155/2012/972629. Epub 2012 Dec 17.


Expression of CD137 on Hodgkin and Reed-Sternberg cells inhibits T-cell activation by eliminating CD137 ligand expression.

Ho WT, Pang WL, Chong SM, Castella A, Al-Salam S, Tan TE, Moh MC, Koh LK, Gan SU, Cheng CK, Schwarz H.

Cancer Res. 2013 Jan 15;73(2):652-61. doi: 10.1158/0008-5472.CAN-12-3849. Epub 2012 Nov 29.


4-1BB ligand-mediated imbalance of helper 17 T cells and regulatory T cells in patients with allergic asthma.

Ai XY, Shi GC, Wan HY, Shi YH, Hou XX, Zhu HX, Tang W.

J Int Med Res. 2012;40(3):1046-54.


Immunotherapy of cancer with 4-1BB.

Vinay DS, Kwon BS.

Mol Cancer Ther. 2012 May;11(5):1062-70. doi: 10.1158/1535-7163.MCT-11-0677. Epub 2012 Apr 24. Review.


Targeting of 4-1BB by monoclonal antibody PF-05082566 enhances T-cell function and promotes anti-tumor activity.

Fisher TS, Kamperschroer C, Oliphant T, Love VA, Lira PD, Doyonnas R, Bergqvist S, Baxi SM, Rohner A, Shen AC, Huang C, Sokolowski SA, Sharp LL.

Cancer Immunol Immunother. 2012 Oct;61(10):1721-33. Epub 2012 Mar 11.


4-1BB ligand modulates direct and Rituximab-induced NK-cell reactivity in chronic lymphocytic leukemia.

Buechele C, Baessler T, Schmiedel BJ, Schumacher CE, Grosse-Hovest L, Rittig K, Salih HR.

Eur J Immunol. 2012 Mar;42(3):737-48. doi: 10.1002/eji.201141920. Epub 2011 Dec 20.


The presence of high level soluble herpes virus entry mediator in sera of gastric cancer patients.

Heo SK, Ju SA, Kim GY, Park SM, Back SH, Park NH, Min YJ, An WG, Nguyen TT, Kim SM, Kim BS.

Exp Mol Med. 2012 Feb 29;44(2):149-58. doi: 10.3858/emm.2012.44.2.010.


Dispensable role for 4-1BB and 4-1BBL in development of vaccinia virus-specific CD8 T cells.

Zhao Y, Croft M.

Immunol Lett. 2012 Jan 30;141(2):220-6. doi: 10.1016/j.imlet.2011.10.008. Epub 2011 Oct 20.


Soluble expression of recombinant human CD137 ligand in Escherichia coli by co-expression of chaperones.

Wang S, Tan A, Lv J, Wang P, Yin X, Chen Y.

J Ind Microbiol Biotechnol. 2012 Mar;39(3):471-6. doi: 10.1007/s10295-011-1045-1. Epub 2011 Oct 15.


CD137 differentially regulates innate and adaptive immunity against Mycobacterium tuberculosis.

Fernández Do Porto DA, Jurado JO, Pasquinelli V, Alvarez IB, Aspera RH, Musella RM, García VE.

Immunol Cell Biol. 2012 Apr;90(4):449-56. doi: 10.1038/icb.2011.63. Epub 2011 Jul 12.

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