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Science. 1977 Feb 11;195(4278):592-4.

Terrestrial locomotion in penguins: it costs more to waddle.

Abstract

The energetic cost for walking is relatively higher for penguins than for other birds or for quadrupeds of similar body mass. The morphology of penguins seems to represent a compromise between aquatic and terrestrial locomotion wherein both energy economy and speed suffer when the birds move on land.

PMID:
835018
[PubMed - indexed for MEDLINE]
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