1 Department of Ecology and Evolutionary Biology, University of California Los Angeles, 621 Charles E. Young Drive South, Los Angeles, CA 90095, USA lawrensack@ucla.edu.
2 IA Watson Grains Research Centre, Plant Breeding Institute, Faculty of Agriculture and Environment, The University of Sydney, Narrabri NSW 2390, Australia.
3 Department of Ecology and Evolutionary Biology, University of California Los Angeles, 621 Charles E. Young Drive South, Los Angeles, CA 90095, USA.