Display Settings:

Format

Send to:

Choose Destination
We are sorry, but NCBI web applications do not support your browser and may not function properly. More information

    PITX2 paired-like homeodomain 2 [ Homo sapiens (human) ]

    Gene ID: 5308, updated on 13-Jun-2013
    Official Symbol
    PITX2provided by HGNC
    Official Full Name
    paired-like homeodomain 2provided by HGNC
    Primary source
    HGNC:9005
    See related
    Ensembl:ENSG00000164093; HPRD:03328; MIM:601542; Vega:OTTHUMG00000132837
    Gene type
    protein coding
    RefSeq status
    REVIEWED
    Organism
    Homo sapiens
    Lineage
    Eukaryota; Metazoa; Chordata; Craniata; Vertebrata; Euteleostomi; Mammalia; Eutheria; Euarchontoglires; Primates; Haplorrhini; Catarrhini; Hominidae; Homo
    Also known as
    RS; RGS; ARP1; Brx1; IDG2; IGDS; IHG2; PTX2; RIEG; IGDS2; IRID2; Otlx2; RIEG1
    Summary
    This gene encodes a member of the RIEG/PITX homeobox family, which is in the bicoid class of homeodomain proteins. The encoded protein acts as a transcription factor and regulates procollagen lysyl hydroxylase gene expression. This protein plays a role in the terminal differentiation of somatotroph and lactotroph cell phenotypes, is involved in the development of the eye, tooth and abdominal organs, and acts as a transcriptional regulator involved in basal and hormone-regulated activity of prolactin. Mutations in this gene are associated with Axenfeld-Rieger syndrome, iridogoniodysgenesis syndrome, and sporadic cases of Peters anomaly. A similar protein in other vertebrates is involved in the determination of left-right asymmetry during development. Alternatively spliced transcript variants encoding distinct isoforms have been described. [provided by RefSeq, Jul 2008]
    Location :
    4q25
    Sequence :
    Chromosome: 4; NC_000004.11 (111538579..111563279, complement)
    See PITX2 in Epigenomics, MapViewer

    Chromosome 4 - NC_000004.11Genomic Context describing neighboring genes Neighboring gene zinc finger protein 969, pseudogene Neighboring gene glutamyl aminopeptidase (aminopeptidase A) Neighboring gene microRNA 297 Neighboring gene lysophospholipase I pseudogene 2

    GeneRIFs: Gene References Into Functions What's a GeneRIF?

    Anophthalmia/Microphthalmia

    Summary from GeneReviews: Anophthalmia / Microphthalmia Overview Go to GeneReviews

    Disease Characteristics
    Anophthalmia refers to complete absence of the globe in the presence of ocular adnexa (eyelids, conjunctiva, and lacrimal apparatus). Microphthalmia is defined as a globe with a total axial length (TAL) that is at least two standard deviations below the mean for age. Classification of microphthalmia is according to the anatomic appearance of the globe and severity of axial length reduction. Severe microphthalmia refers to a globe with a corneal diameter less than 4 mm and a TAL less than 10 mm at birth or less than 12 mm after age one year. Simple microphthalmia refers to an eye that is anatomically intact except for its short TAL. Complex microphthalmia refers to an eye with anterior segment dysgenesis and/or posterior segment dysgenesis. Anophthalmia/microphthalmia (A/M) can be unilateral or bilateral. A/M is a heterogenous condition with various etiologies. It can be isolated or can occur with other anomalies or as part of a well-defined syndrome. One-third of individuals with A/M have associated malformations. Heritable causes of A/M include chromosome abnormalities and syndromic or nonsyndromic single gene disorders.
    Diagnosis Testing
    The diagnosis of anophthalmia/microphthalmia (A/M) is based on clinical examination and imaging studies including: A-scan ultrasonography to measure total axial length; B-scan ultrasonography to evaluate the internal structures of the globe; and CT scan or MRI of the brain and orbits to evaluate the size and internal structures of the globe, the optic nerve and extraocular muscles, and brain anatomy. Evaluation for other malformations, assessment of hearing, chromosome analysis, family history, and parental eye examinations may help establish the underlying cause. Mutations in the following genes are associated with A/M: SIX3, HESX1, BCOR, SHH, PAX6, RAX, CHD7 (CHARGE syndrome), IKBKG (incontinentia pigmenti), NDP (Norrie disease), SOX2 (SOX2-related eye disorders), POMT1 (Walker-Warburg syndrome), and SIX6.
    Genetic Counseling
    If a proband has an inherited or de novo chromosome abnormality or a specific syndrome associated with A/M, genetic counseling, and possibly genetic testing, for that condition is indicated. Rarely, isolated anophthalmia may be inherited in an autosomal dominant, autosomal recessive, or X-linked manner. Assuming that one-half of cases are sporadic and one-half inherited, the empiric risk for sibs without a clear etiology or family history is 10%-15%. Prenatal diagnosis for pregnancies at increased risk for chromosome abnormalities and some syndromic forms of A/M is possible. Transvaginal ultrasound examination may detect the eyes from 12 weeks' gestation onward; in most reports, eye malformations are detected only after 22 weeks' gestation. The sensitivity of transvaginal ultrasound examination in detecting A/M is not known. Three-dimensional and four-dimensional ultrasound examination may be used in some centers to detect complex malformations of the face, including A/M. MRI may be a useful adjunct to ultrasound examination in detection of anophthalmia.
    References
    Products Interactant Other Gene Complex Source Pubs Description
    Q99697 P35222 CTNNB1    HPRD  PubMed  
    Q99697 Q13547 HDAC1    HPRD  PubMed  
    Q99697 Q92993 KAT5    HPRD  PubMed  
    Q99697 Q9UJU2 LEF1    HPRD  PubMed  
    Q99697 O00151 PDLIM1    HPRD  PubMed  
    Q99697 Q99697 PITX2    HPRD  PubMed  
    Q99697 O75360 PROP1    HPRD  PubMed  
    BioGRID:111325 BioGRID:107880 CTNNB1    BioGRID  PubMed Affinity Capture-Western 
    BioGRID:111325 BioGRID:109315 HDAC1    BioGRID  PubMed Affinity Capture-Western 
    BioGRID:111325 BioGRID:119365 HERC5    BioGRID  PubMed Reconstituted Complex 
    BioGRID:111325 BioGRID:109438 HOXA1    BioGRID  PubMed Affinity Capture-Western; PCA 
    BioGRID:111325 BioGRID:200362 Hoxa1    BioGRID  PubMed Two-hybrid 
    BioGRID:111325 BioGRID:110594 MSX2    BioGRID  PubMed Phenotypic Suppression 
    BioGRID:111325 BioGRID:111325 PITX2    BioGRID  PubMed Affinity Capture-Western; Reconstituted Complex 
    BioGRID:111325 BioGRID:247549 Pou1f1    BioGRID  PubMed Phenotypic Enhancement; Reconstituted Complex 
    BioGRID:111325 BioGRID:110263 SMAD3    BioGRID  PubMed Affinity Capture-Western 
    BioGRID:111325 BioGRID:113500 TRIM25    BioGRID  PubMed Reconstituted Complex; Two-hybrid 
    BioGRID:111325 BioGRID:116272 WDR5    BioGRID  PubMed Affinity Capture-Western 
    BioGRID:111325 BioGRID:114736 ZNHIT3    BioGRID  PubMed Two-hybrid 
    • Heart Development, organism-specific biosystem (from WikiPathways)
      Heart Development, organism-specific biosystemThis pathway has been largely adapted from an article by Deepak Srivastava, Cell. 2006 Sep 22;126(6):1037-48. In this pathway are known transcription factors, miRNAs and regulatory proteins that impa...
    • Integrated Pancreatic Cancer Pathway, organism-specific biosystem (from WikiPathways)
      Integrated Pancreatic Cancer Pathway, organism-specific biosystemAn integrated pathway model which displays the protein-protein interactions (PPIs) among the relevant proteins for pancreatic cancer. This pathway is a collection of different mechanistic protein pat...
    • Regulation of Wnt-mediated beta catenin signaling and target gene transcription, organism-specific biosystem (from Pathway Interaction Database)
      Regulation of Wnt-mediated beta catenin signaling and target gene transcription, organism-specific biosystem
      Regulation of Wnt-mediated beta catenin signaling and target gene transcription
    • TGF-beta signaling pathway, organism-specific biosystem (from KEGG)
      TGF-beta signaling pathway, organism-specific biosystemThe transforming growth factor-beta (TGF-beta) family members, which include TGF-betas, activins and bone morphogenetic proteins (BMPs), are structurally related secreted cytokines found in species r...
    • TGF-beta signaling pathway, conserved biosystem (from KEGG)
      TGF-beta signaling pathway, conserved biosystemThe transforming growth factor-beta (TGF-beta) family members, which include TGF-betas, activins and bone morphogenetic proteins (BMPs), are structurally related secreted cytokines found in species r...

    Markers

    Homology

    Clone Names

    • MGC20144, MGC111022

    Gene Ontology Provided by GOA

    Function Evidence Code Pubs
    RNA polymerase II activating transcription factor binding IPI
    Inferred from Physical Interaction
    more info
    PubMed 
    RNA polymerase II core promoter proximal region sequence-specific DNA binding IEA
    Inferred from Electronic Annotation
    more info
     
    RNA polymerase II core promoter proximal region sequence-specific DNA binding transcription factor activity involved in negative regulation of transcription IDA
    Inferred from Direct Assay
    more info
    PubMed 
    RNA polymerase II core promoter proximal region sequence-specific DNA binding transcription factor activity involved in positive regulation of transcription IEA
    Inferred from Electronic Annotation
    more info
     
    RNA polymerase II transcription coactivator activity IDA
    Inferred from Direct Assay
    more info
    PubMed 
    RNA polymerase II transcription factor binding IPI
    Inferred from Physical Interaction
    more info
    PubMed 
    RNA polymerase II transcription factor binding transcription factor activity involved in negative regulation of transcription IDA
    Inferred from Direct Assay
    more info
    PubMed 
    chromatin DNA binding IEA
    Inferred from Electronic Annotation
    more info
     
    phosphoprotein binding IPI
    Inferred from Physical Interaction
    more info
    PubMed 
    protein binding IPI
    Inferred from Physical Interaction
    more info
    PubMed 
    protein homodimerization activity IPI
    Inferred from Physical Interaction
    more info
    PubMed 
    ribonucleoprotein complex binding IC
    Inferred by Curator
    more info
    PubMed 
    sequence-specific DNA binding transcription factor activity IDA
    Inferred from Direct Assay
    more info
    PubMed 
    transcription factor binding IPI
    Inferred from Physical Interaction
    more info
    PubMed 
    transcription regulatory region sequence-specific DNA binding IDA
    Inferred from Direct Assay
    more info
    PubMed 
    Process Evidence Code Pubs
    Wnt receptor signaling pathway IEA
    Inferred from Electronic Annotation
    more info
     
    atrial cardiac muscle tissue morphogenesis IEA
    Inferred from Electronic Annotation
    more info
     
    atrioventricular valve development IEA
    Inferred from Electronic Annotation
    more info
     
    camera-type eye development IMP
    Inferred from Mutant Phenotype
    more info
    PubMed 
    cardiac neural crest cell migration involved in outflow tract morphogenesis ISS
    Inferred from Sequence or Structural Similarity
    more info
     
    cell proliferation involved in outflow tract morphogenesis ISS
    Inferred from Sequence or Structural Similarity
    more info
     
    deltoid tuberosity development IMP
    Inferred from Mutant Phenotype
    more info
    PubMed 
    determination of left/right symmetry ISS
    Inferred from Sequence or Structural Similarity
    more info
     
    embryonic camera-type eye development IEA
    Inferred from Electronic Annotation
    more info
     
    embryonic digestive tract morphogenesis IEA
    Inferred from Electronic Annotation
    more info
     
    embryonic hindlimb morphogenesis IEA
    Inferred from Electronic Annotation
    more info
     
    endodermal digestive tract morphogenesis IEA
    Inferred from Electronic Annotation
    more info
     
    extraocular skeletal muscle development IEA
    Inferred from Electronic Annotation
    more info
     
    female gonad development IEA
    Inferred from Electronic Annotation
    more info
     
    hair cell differentiation IC
    Inferred by Curator
    more info
    PubMed 
    hypothalamus cell migration IEA
    Inferred from Electronic Annotation
    more info
     
    in utero embryonic development IEA
    Inferred from Electronic Annotation
    more info
     
    iris morphogenesis IMP
    Inferred from Mutant Phenotype
    more info
    PubMed 
    left lung morphogenesis IEA
    Inferred from Electronic Annotation
    more info
     
    left/right axis specification ISS
    Inferred from Sequence or Structural Similarity
    more info
     
    male gonad development IEA
    Inferred from Electronic Annotation
    more info
     
    myoblast fusion IEA
    Inferred from Electronic Annotation
    more info
     
    negative regulation of transcription from RNA polymerase II promoter IDA
    Inferred from Direct Assay
    more info
    PubMed 
    neuron migration IEA
    Inferred from Electronic Annotation
    more info
     
    odontogenesis IMP
    Inferred from Mutant Phenotype
    more info
    PubMed 
    odontogenesis of dentin-containing tooth IEA
    Inferred from Electronic Annotation
    more info
     
    patterning of blood vessels IEA
    Inferred from Electronic Annotation
    more info
     
    positive regulation of DNA binding IEA
    Inferred from Electronic Annotation
    more info
     
    positive regulation of myoblast proliferation IEA
    Inferred from Electronic Annotation
    more info
     
    positive regulation of transcription from RNA polymerase II promoter IDA
    Inferred from Direct Assay
    more info
    PubMed 
    prolactin secreting cell differentiation TAS
    Traceable Author Statement
    more info
    PubMed 
    pulmonary myocardium development IEA
    Inferred from Electronic Annotation
    more info
     
    pulmonary vein morphogenesis IEA
    Inferred from Electronic Annotation
    more info
     
    regulation of cell migration IEA
    Inferred from Electronic Annotation
    more info
     
    regulation of transcription, DNA-dependent IDA
    Inferred from Direct Assay
    more info
    PubMed 
    response to hormone stimulus IEA
    Inferred from Electronic Annotation
    more info
     
    response to vitamin A IEA
    Inferred from Electronic Annotation
    more info
     
    somatotropin secreting cell differentiation TAS
    Traceable Author Statement
    more info
    PubMed 
    spleen development ISS
    Inferred from Sequence or Structural Similarity
    more info
     
    subthalamic nucleus development IEA
    Inferred from Electronic Annotation
    more info
     
    superior vena cava morphogenesis IEA
    Inferred from Electronic Annotation
    more info
     
    vascular smooth muscle cell differentiation IEA
    Inferred from Electronic Annotation
    more info
     
    vasculogenesis IEA
    Inferred from Electronic Annotation
    more info
     
    ventricular cardiac muscle cell development IEA
    Inferred from Electronic Annotation
    more info
     
    ventricular septum morphogenesis IEA
    Inferred from Electronic Annotation
    more info
     
    Component Evidence Code Pubs
    cytoplasm IEA
    Inferred from Electronic Annotation
    more info
     
    nucleus IDA
    Inferred from Direct Assay
    more info
    PubMed 
    transcription factor complex IDA
    Inferred from Direct Assay
    more info
    PubMed 
    Preferred Names
    pituitary homeobox 2
    Names
    pituitary homeobox 2
    solurshin
    all1-responsive gene 1
    homeobox protein PITX2
    ALL1-responsive protein ARP1
    paired-like homeodomain transcription factor 2
    rieg bicoid-related homeobox transcription factor 1

    RefSeqs maintained independently of Annotated Genomes

    These reference sequences exist independently of genome builds. Explain

    These reference sequences are curated independently of the genome annotation cycle, so their versions may not match the RefSeq versions in the current genome build. Identify version mismatches by comparing the version of the RefSeq in this section to the one reported in Genomic regions, transcripts, and products above.

    Genomic

    1. NG_007120.1 RefSeqGene

      Range
      230..24930
      Download
      GenBank, FASTA, Sequence Viewer (Graphics)

    mRNA and Protein(s)

    1. NM_000325.5NP_000316.2  pituitary homeobox 2 isoform c

      Status: REVIEWED

      Description
      Transcript Variant: This variant (3), also known as ARP1c, lacks several exons at the 5' end and has an alternate 5' exon, as compared to variant 2. The resulting isoform (c) has a longer and distinct N-terminus, as compared to isoform b.
      Source sequence(s)
      AC017068, BC013998
      Consensus CDS
      CCDS3694.1
      UniProtKB/Swiss-Prot
      Q99697
      Related
      ENSP00000304169, OTTHUMP00000163738, ENST00000306732, OTTHUMT00000256310
      Conserved Domains (3) summary
      cd00086
      Location:93151
      Blast Score: 196
      homeodomain; Homeodomain; DNA binding domains involved in the transcriptional regulation of key eukaryotic developmental processes; may bind to DNA as monomers or as homo- and/or heterodimers, in a sequence-specific manner.
      COG5576
      Location:40160
      Blast Score: 138
      COG5576; Homeodomain-containing transcription factor [Transcription]
      pfam03826
      Location:281299
      Blast Score: 94
      OAR; OAR domain
    2. NM_001204397.1NP_001191326.1  pituitary homeobox 2 isoform b

      Status: REVIEWED

      Description
      Transcript Variant: This variant (4) has an additional exon in the 5' UTR, as compared to variant 2. Variants 2, 4 and 5 encode the same isoform b.
      Source sequence(s)
      AK127829, BC106010, BP372081, BX380370, U69961
      Consensus CDS
      CCDS3692.1
      UniProtKB/Swiss-Prot
      Q99697
      Related
      ENSP00000347004, OTTHUMP00000163736, ENST00000354925, OTTHUMT00000256308
      Conserved Domains (2) summary
      cd00086
      Location:86144
      Blast Score: 189
      homeodomain; Homeodomain; DNA binding domains involved in the transcriptional regulation of key eukaryotic developmental processes; may bind to DNA as monomers or as homo- and/or heterodimers, in a sequence-specific manner.
      pfam03826
      Location:274292
      Blast Score: 98
      OAR; OAR domain
    3. NM_001204398.1NP_001191327.1  pituitary homeobox 2 isoform b

      Status: REVIEWED

      Description
      Transcript Variant: This variant (5) lacks an exon in the 5' UTR, as compared to variant 2. Variants 2, 4 and 5 encode the same isoform b.
      Source sequence(s)
      BC106010, BP372081
      Consensus CDS
      CCDS3692.1
      UniProtKB/Swiss-Prot
      Q99697
      Conserved Domains (2) summary
      cd00086
      Location:86144
      Blast Score: 189
      homeodomain; Homeodomain; DNA binding domains involved in the transcriptional regulation of key eukaryotic developmental processes; may bind to DNA as monomers or as homo- and/or heterodimers, in a sequence-specific manner.
      pfam03826
      Location:274292
      Blast Score: 98
      OAR; OAR domain
    4. NM_001204399.1NP_001191328.1  pituitary homeobox 2 isoform a

      Status: REVIEWED

      Description
      Transcript Variant: This variant (6) lacks an exon in the 5' UTR and an in-frame exon in the 5' CDS, as compared to variant 2. The resulting isoform (a) lacks an internal segment, as compared to isoform b. Variants 1 and 6 encode the same isoform a.
      Source sequence(s)
      BC106010, BP372081
      Consensus CDS
      CCDS3693.1
      UniProtKB/Swiss-Prot
      Q99697
      Conserved Domains (2) summary
      pfam03826
      Location:228246
      Blast Score: 99
      OAR; OAR domain
      cd00086
      Location:4098
      Blast Score: 214
      homeodomain; Homeodomain; DNA binding domains involved in the transcriptional regulation of key eukaryotic developmental processes; may bind to DNA as monomers or as homo- and/or heterodimers, in a sequence-specific manner.
    5. NM_153426.2NP_700475.1  pituitary homeobox 2 isoform b

      Status: REVIEWED

      Description
      Transcript Variant: This variant (2), also known as ARP1b, encodes the predominant isoform (b).
      Source sequence(s)
      AK127829, BC013998, BC106010, BP372081, BX380370, U69961
      Consensus CDS
      CCDS3692.1
      UniProtKB/Swiss-Prot
      Q99697
      Conserved Domains (2) summary
      cd00086
      Location:86144
      Blast Score: 189
      homeodomain; Homeodomain; DNA binding domains involved in the transcriptional regulation of key eukaryotic developmental processes; may bind to DNA as monomers or as homo- and/or heterodimers, in a sequence-specific manner.
      pfam03826
      Location:274292
      Blast Score: 98
      OAR; OAR domain
    6. NM_153427.2NP_700476.1  pituitary homeobox 2 isoform a

      Status: REVIEWED

      Description
      Transcript Variant: This variant (1), also known as ARP1a, lacks an in-frame exon in the 5' region, as compared to variant 2. The resulting isoform (a) lacks an internal segment, as compared to isoform b. Variants 1 and 6 encode the same isoform a.
      Source sequence(s)
      AK127829, BC106010, BP372081, BX380370, U69961
      Consensus CDS
      CCDS3693.1
      UniProtKB/Swiss-Prot
      Q99697
      Conserved Domains (2) summary
      pfam03826
      Location:228246
      Blast Score: 99
      OAR; OAR domain
      cd00086
      Location:4098
      Blast Score: 214
      homeodomain; Homeodomain; DNA binding domains involved in the transcriptional regulation of key eukaryotic developmental processes; may bind to DNA as monomers or as homo- and/or heterodimers, in a sequence-specific manner.

    RefSeqs of Annotated Genomes: Homo sapiens Annotation Release 104

    The following sections contain reference sequences that belong to a specific genome build. Explain

    Reference GRCh37.p10 Primary Assembly

    Genomic

    1. NC_000004.11 Reference GRCh37.p10 Primary Assembly

      Range
      111538579..111563279, complement
      Download
      GenBank, FASTA, Sequence Viewer (Graphics)

    Alternate HuRef

    Genomic

    1. AC_000136.1 Alternate HuRef

      Range
      107270120..107294817, complement
      Download
      GenBank, FASTA, Sequence Viewer (Graphics)

    Alternate CHM1_1.0

    Genomic

    1. NC_018915.1 Alternate CHM1_1.0

      Range
      111332856..111357552, complement
      Download
      GenBank, FASTA, Sequence Viewer (Graphics)

      Supplemental Content

      Recent activity

      Your browsing activity is empty.

      Activity recording is turned off.

      Turn recording back on

      See more...