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    NPC1 Niemann-Pick disease, type C1 [ Homo sapiens ]

    Gene ID: 4864, updated on 13-May-2012

    Summary

    Official Symbol
    NPC1provided by HGNC
    Official Full Name
    Niemann-Pick disease, type C1provided by HGNC
    Primary source
    HGNC:7897
    See related
    Ensembl:ENSG00000141458; HPRD:09622; MIM:607623; Vega:OTTHUMG00000131873
    Gene type
    protein coding
    RefSeq status
    REVIEWED
    Organism
    Homo sapiens
    Lineage
    Eukaryota; Metazoa; Chordata; Craniata; Vertebrata; Euteleostomi; Mammalia; Eutheria; Euarchontoglires; Primates; Haplorrhini; Catarrhini; Hominidae; Homo
    Also known as
    NPC; FLJ98532
    Summary
    This gene encodes a large protein that resides in the limiting membrane of endosomes and lysosomes and mediates intracellular cholesterol trafficking via binding of cholesterol to its N-terminal domain. It is predicted to have a cytoplasmic C-terminus, 13 transmembrane domains, and 3 large loops in the lumen of the endosome - the last loop being at the N-terminus. This protein transports low-density lipoproteins to late endosomal/lysosomal compartments where they are hydrolized and released as free cholesterol. Defects in this gene cause Niemann-Pick type C disease, a rare autosomal recessive neurodegenerative disorder characterized by over accumulation of cholesterol and glycosphingolipids in late endosomal/lysosomal compartments.[provided by RefSeq, Aug 2009]

    Genomic context

    Location :
    18q11-q12
    Sequence :
    Chromosome: 18; NC_000018.9 (21111463..21166581, complement)
    See NPC1 in Epigenomics, MapViewer

    Chromosome 18 - NC_000018.9Genomic Context describing neighboring genes Neighboring gene transmembrane protein 241 Neighboring gene RIO kinase 3 (yeast) Neighboring gene chromosome 18 open reading frame 8 Neighboring gene ankyrin repeat domain 29 Neighboring gene ribosomal protein S10 pseudogene 27

    Genomic regions, transcripts, and products

    Bibliography

    GeneRIFs: Gene References Into Functions What's a GeneRIF?

    Phenotypes

    Genome-wide association study for early-onset and morbid adult obesity identifies three new risk loci in European populations.

    Niemann-Pick disease, type C1

    Summary from GeneReviews: Go to GeneReviews

    Disease Characteristics
    Niemann-Pick disease type C (NPC) is a lipid storage disease that can present in infants, children, or adults. Neonates can present with ascites and severe liver disease from infiltration of the liver and/or respiratory failure from infiltration of the lungs. Other infants, without liver or pulmonary disease, have hypotonia and developmental delay. The classic presentation occurs in mid-to-late childhood with the insidious onset of ataxia, vertical supranuclear gaze palsy (VSGP), and dementia. Dystonia and seizures are common. Dysarthria and dysphagia eventually become disabling, making oral feeding impossible; death usually occurs in the late second or third decade from aspiration pneumonia. Adults are more likely to present with dementia or psychiatric symptoms.
    Diagnosis Testing
    The diagnosis of NPC is confirmed by biochemical testing that demonstrates impaired cholesterol esterification and positive filipin staining in cultured fibroblasts. Biochemical testing for carrier status is unreliable. Most individuals with NPC have NPC1, caused by mutations in the NPC1 gene; fewer than 20 individuals have been diagnosed with NPC2, caused by mutations in the NPC2 gene. Molecular genetic testing of the NPC1 and NPC2 genes detects disease-causing mutations in approximately 94% of individuals with NPC. Such testing is available clinically.
    Genetic Counseling
    NPC is inherited in an autosomal recessive manner. Each sib of an affected individual has a 25% chance of being affected, a 50% chance of being an asymptomatic carrier, and a 25% chance of being unaffected and not a carrier. The phenotype (i.e., age of onset and severity of symptoms) usually runs true in families. Carrier testing for at-risk relatives and prenatal testing for pregnancies at increased risk are possible when the two disease-causing mutations have been identified in the family.
    References

    Interactions

    Products Interactant Other Gene Complex Source Pubs Description
    BioGRID:110925 BioGRID:110219 LRP6    BioGRID  PubMed Affinity Capture-Western 
    BioGRID:110925 BioGRID:125380 OSBPL5    BioGRID  PubMed Affinity Capture-Western 
    BioGRID:110925 BioGRID:130988 TMEM173    BioGRID  PubMed Affinity Capture-MS 
    BioGRID:110925 BioGRID:113164 UBC    BioGRID  PubMed Affinity Capture-MS; Affinity Capture-Western 
    BioGRID:110925 BioGRID:123557 UNC93B1    BioGRID  PubMed Affinity Capture-MS 
    BioGRID:110925 BioGRID:118059 VPS4A    BioGRID  PubMed Affinity Capture-Western; Co-localization; Reconstituted Complex 

    General gene information

    Markers

    Homology

    • Homologs of the NPC1 gene: The NPC1 gene is conserved in chimpanzee, , dog, cow, mouse, rat, chicken, zebrafish, fruit fly, mosquito, S.cerevisiae, K.lactis, , , N.crassa, and A.thaliana.
    • Map Viewer (Mouse, Rat)

    Pathways from BioSystems

    • Lysosome, organism-specific biosystem (from KEGG)
      Lysosome, organism-specific biosystemLysosomes are membrane-delimited organelles in animal cells serving as the cell's main digestive compartment to which all sorts of macromolecules are delivered for degradation. They contain more than...
    • Lysosome, conserved biosystem (from KEGG)
      Lysosome, conserved biosystemLysosomes are membrane-delimited organelles in animal cells serving as the cell's main digestive compartment to which all sorts of macromolecules are delivered for degradation. They contain more than...

    Gene Ontology Provided by GOA

    Function Evidence Code Pubs
    hedgehog receptor activity IEA
    Inferred from Electronic Annotation
    more info
     
    protein binding IPI
    Inferred from Physical Interaction
    more info
    PubMed 
    receptor activity TAS
    Traceable Author Statement
    more info
    PubMed 
    sterol transporter activity TAS
    Traceable Author Statement
    more info
    PubMed 
    transmembrane signaling receptor activity TAS
    Traceable Author Statement
    more info
    PubMed 
    Process Evidence Code Pubs
    autophagy IGI
    Inferred from Genetic Interaction
    more info
    PubMed 
    bile acid metabolic process ISS
    Inferred from Sequence or Structural Similarity
    more info
     
    cholesterol efflux IDA
    Inferred from Direct Assay
    more info
    PubMed 
    cholesterol homeostasis IDA
    Inferred from Direct Assay
    more info
    PubMed 
    cholesterol homeostasis ISS
    Inferred from Sequence or Structural Similarity
    more info
     
    lysosomal transport ISS
    Inferred from Sequence or Structural Similarity
    more info
     
    sterol transport TAS
    Traceable Author Statement
    more info
    PubMed 
    Component Evidence Code Pubs
    endoplasmic reticulum IDA
    Inferred from Direct Assay
    more info
    PubMed 
    endosome IEA
    Inferred from Electronic Annotation
    more info
     
    integral to membrane TAS
    Traceable Author Statement
    more info
    PubMed 
    integral to plasma membrane IDA
    Inferred from Direct Assay
    more info
    PubMed 
    late endosome membrane IEA
    Inferred from Electronic Annotation
    more info
     
    lysosomal membrane IEA
    Inferred from Electronic Annotation
    more info
     
    lysosome TAS
    Traceable Author Statement
    more info
    PubMed 
    membrane IEA
    Inferred from Electronic Annotation
    more info
     
    nuclear envelope IDA
    Inferred from Direct Assay
    more info
    PubMed 
    perinuclear region of cytoplasm IDA
    Inferred from Direct Assay
    more info
    PubMed 

    General protein information

    Preferred Names
    Niemann-Pick C1 protein
    Names
    Niemann-Pick C1 protein

    NCBI Reference Sequences (RefSeq)

    RefSeqs maintained independently of Annotated Genomes

    These reference sequences exist independently of genome builds. Explain

    These reference sequences are curated independently of the genome annotation cycle, so their versions may not match the RefSeq versions in the current genome build. Identify version mismatches by comparing the version of the RefSeq in this section to the one reported in Genomic regions, transcripts, and products above.

    Genomic

    1. NG_012795.1 RefSeqGene

      Range
      5001..60119
      Download
      GenBank, FASTA, Sequence Viewer (Graphics)

    mRNA and Protein(s)

    1. NM_000271.4NP_000262.2  Niemann-Pick C1 protein precursor

      Status: REVIEWED

      Source sequence(s)
      AB209048, AF002020, BC063302, DR159421
      Consensus CDS
      CCDS11878.1
      UniProtKB/Swiss-Prot
      O15118
      Related
      ENSP00000269228, OTTHUMP00000162699, ENST00000269228, OTTHUMT00000254823
      Conserved Domains (2) summary
      TIGR00917
      Location:251252
      Blast Score: 4868
      2A060601; Niemann-Pick C type protein family
      cl14618
      Location:649802
      Blast Score: 540
      SecD_SecF; Protein export membrane protein

    RefSeqs of Annotated Genomes: Build 37.3

    The following sections contain reference sequences that belong to a specific genome build. Explain

    Reference GRCh37.p5 Primary Assembly

    Genomic

    1. NC_000018.9 Reference GRCh37.p5 Primary Assembly

      Range
      21111463..21166581, complement
      Download
      GenBank, FASTA, Sequence Viewer (Graphics)

    Alternate HuRef

    Genomic

    1. AC_000150.1 Alternate HuRef

      Range
      17966414..18021712, complement
      Download
      GenBank, FASTA, Sequence Viewer (Graphics)

    Related Sequences

    Nucleotide Protein
    Heading Accession and Version
    genomic AC010853.11 (84944..139909) None
    genomic AF157379.1 AAD48006.1
    genomic AF338230.2 AAK25791.1
    genomic CH471088.1 EAX01158.1
    mRNA AF002020.1 AAB63982.1
    mRNA AI242474.1 None
    mRNA AK293779.1 BAG57194.1
    mRNA AK295715.1 BAG58559.1
    mRNA AK308491.1 None
    mRNA BC063302.1 AAH63302.1
    mRNA BP217319.1 None
    mRNA CA438246.1 None
    mRNA DR159421.1 None
    mRNA AB209048.1 BAD92285.1
    Protein Accession Links
    GenPept Link UniProtKB Link
    O15118.2 GenPept UniProtKB/Swiss-Prot:O15118
    Q59GR1 GenPept UniProtKB/TrEMBL:Q59GR1

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