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    SH2D1A SH2 domain containing 1A [ Homo sapiens (human) ]

    Gene ID: 4068, updated on 5-May-2013
    Official Symbol
    SH2D1Aprovided by HGNC
    Official Full Name
    SH2 domain containing 1Aprovided by HGNC
    Primary source
    HGNC:10820
    Locus tag
    RP5-1052M9.3
    See related
    Ensembl:ENSG00000183918; HPRD:02390; MIM:300490; Vega:OTTHUMG00000022344
    Gene type
    protein coding
    RefSeq status
    REVIEWED
    Organism
    Homo sapiens
    Lineage
    Eukaryota; Metazoa; Chordata; Craniata; Vertebrata; Euteleostomi; Mammalia; Eutheria; Euarchontoglires; Primates; Haplorrhini; Catarrhini; Hominidae; Homo
    Also known as
    LYP; SAP; XLP; DSHP; EBVS; IMD5; XLPD; MTCP1; SAP/SH2D1A
    Summary
    This gene encodes a protein that plays a major role in the bidirectional stimulation of T and B cells. This protein contains an SH2 domain and a short tail. It associates with the signaling lymphocyte-activation molecule, thereby acting as an inhibitor of this transmembrane protein by blocking the recruitment of the SH2-domain-containing signal-transduction molecule SHP-2 to its docking site. This protein can also bind to other related surface molecules that are expressed on activated T, B and NK cells, thereby modifying signal transduction pathways in these cells. Mutations in this gene cause lymphoproliferative syndrome X-linked type 1 or Duncan disease, a rare immunodeficiency characterized by extreme susceptibility to infection with Epstein-Barr virus, with symptoms including severe mononucleosis and malignant lymphoma. Multiple transcript variants encoding different isoforms have been found for this gene. [provided by RefSeq, Jul 2008]
    Location :
    Xq25
    Sequence :
    Chromosome: X; NC_000023.10 (123480132..123507010)
    See SH2D1A in Epigenomics, MapViewer

    Chromosome X - NC_000023.10Genomic Context describing neighboring genes Neighboring gene testis expressed sequence 13-like pseudogene Neighboring gene nucleophosmin 1 (nucleolar phosphoprotein B23, numatrin) pseudogene 34 Neighboring gene teneurin transmembrane protein 1 Neighboring gene ribosomal protein S26 pseudogene 57 Neighboring gene testis expressed sequence 13-like

    GeneRIFs: Gene References Into Functions What's a GeneRIF?

    X-linked lymphoproliferative syndrome 1

    Summary from GeneReviews: Lymphoproliferative Disease, X-Linked Go to GeneReviews

    Disease Characteristics
    X-linked lymphoproliferative disease (XLP) is caused by mutations in SH2D1A and XIAP (BIRC4). XLP may also occur in rare instances with no identified underlying genetic cause. The three most commonly recognized phenotypes of SH2D1A-related XLP are hemophagocytic lymphohistiocytosis (HLH) associated with Epstein-Barr virus (EBV) infection (58% of individuals), dysgammaglobulinemia (31%), and lymphoproliferative disorders (malignant lymphoma) (30%). Manifestations of SH2D1A-related XLP, including HLH, can also occur in the absence of EBV. XIAP-related XLP also presents with HLH (often associated with EBV) or dysgammaglobulinemia, but no cases of lymphoma have been described to date. HLH resulting from EBV infection, sometimes referred to as severe infectious mononucleosis, is associated with an unregulated and exaggerated immune response with widespread proliferation of cytotoxic T cells, EBV-infected B cells, and macrophages. Fulminant hepatitis, hepatic necrosis, and profound bone marrow failure are typical, resulting in mortality that is higher than 90%, though prompt recognition of the disorder and aggressive treatment interventions likely improve survival. Dysgammaglobulinemia is typically hypogammaglobulinemia of one or more immunoglobulin subclasses. The prognosis is improved if affected males are managed with regular intravenous immunoglobulin (IVIG) therapy. The malignant lymphomas are typically high-grade B cell lymphomas, non-Hodgkin type, often extranodal, and in particular involve the intestine.
    Diagnosis Testing
    XLP is typically suspected based on clinical history and tests of immune function. Diagnosis can be confirmed with molecular genetic testing of SH2D1A and/or XIAP, which is available on a clinical basis. SH2D1A and XIAP encode SLAM-associated protein (SAP) and baculoviral IAP repeat-containing protein 4 (X-linked inhibitor of apoptosis; XIAP), respectively. Absence of SAP or XIAP strongly supports the diagnosis of XLP as well.
    Genetic Counseling
    XLP is inherited in an X-linked manner. Carrier females have a 50% chance of transmitting the disease-causing mutation in each pregnancy: males who inherit the mutation will be affected; females who inherit the mutation will be carriers. Carrier testing of at-risk female relatives and prenatal testing for pregnancies at increased risk are possible if the disease-causing mutation in the family is known. Because of the possibility of germline mosaicism, it is appropriate to offer prenatal diagnosis to couples who have had a child with XLP if the disease-causing mutation in the proband is known.
    References
    Products Interactant Other Gene Complex Source Pubs Description
    NP_002342.1 NP_057466.1 CD244    BIND  PubMed SH2D1A interacts with CD244. 
    NP_002342.1 NP_003865.1 CD84    BIND  PubMed SH2D1A interacts with CD84. 
    NP_002342.1 NP_002339.1 LY9    BIND  PubMed SH2D1A interacts with CD229. 
    NP_002342.1 NP_003028.1 SLAMF1    BIND  PubMed SLAM interacts with SAP in a phosphorylation-independent manner. 
    NP_002342.1 NP_003028.1 SLAMF1    BIND  PubMed SH2D1A interacts with CD150. 
    O60880 Q15052 ARHGEF6    HPRD  PubMed  
    O60880 Q14155 ARHGEF7    HPRD  PubMed  
    O60880 Q9UIB8 CD84    HPRD  PubMed  
    O60880 Q99704 DOK1    HPRD  PubMed  
    O60880 O15117 FYB    HPRD  PubMed  
    O60880 P06241 FYN    HPRD  PubMed  
    O60880 P62993 GRB2    HPRD  PubMed  
    O60880 P06239 LCK    HPRD  PubMed  
    O60880 Q9HBG7 LY9    HPRD  PubMed  
    O60880 Q13291 SLAMF1    HPRD  PubMed  
    O60880 Q96DU3 SLAMF6    HPRD  PubMed  
    O60880 Q9NQ25 SLAMF7    HPRD  PubMed  
    BioGRID:110246 BioGRID:119709 CD244    BioGRID  PubMed Affinity Capture-Western 
    BioGRID:110246 BioGRID:114359 CD84    BioGRID  PubMed Affinity Capture-Western 
    BioGRID:110246 BioGRID:108131 DOK1    BioGRID  PubMed Affinity Capture-Western; Reconstituted Complex 
    BioGRID:110246 BioGRID:108276 EGFR    BioGRID  PubMed Protein-peptide 
    BioGRID:110246 BioGRID:108309 ELAVL1    BioGRID  PubMed Affinity Capture-RNA 
    BioGRID:110246 BioGRID:108377 ERBB3    BioGRID  PubMed Protein-peptide 
    BioGRID:110246 BioGRID:108810 FYN    BioGRID  PubMed Affinity Capture-Western; Reconstituted Complex 
    BioGRID:110246 BioGRID:110241 LY9    BioGRID  PubMed Affinity Capture-Western; Two-hybrid 
    BioGRID:110246 BioGRID:111580 MAPK1    BioGRID  PubMed Biochemical Activity 
    BioGRID:110246 BioGRID:111586 MAPK11    BioGRID  PubMed Biochemical Activity 
    BioGRID:110246 BioGRID:107819 MAPK14    BioGRID  PubMed Biochemical Activity 
    BioGRID:110246 BioGRID:112395 SLAMF1    BioGRID  PubMed Affinity Capture-Western; Reconstituted Complex; Two-hybrid 
    BioGRID:110246 BioGRID:125378 SLAMF6    BioGRID  PubMed Affinity Capture-Western 
    BioGRID:110246 BioGRID:113164 UBC    BioGRID  PubMed Affinity Capture-MS 
    • Direct p53 effectors, organism-specific biosystem (from Pathway Interaction Database)
      Direct p53 effectors, organism-specific biosystem
      Direct p53 effectors
    • Measles, organism-specific biosystem (from KEGG)
      Measles, organism-specific biosystemMeasles virus (MV) is highly contagious virus that leads infant death worldwide. Humans are the unique natural reservoir for this virus. It causes severe immunosuppression favouring secondary bacteri...
    • Measles, conserved biosystem (from KEGG)
      Measles, conserved biosystemMeasles virus (MV) is highly contagious virus that leads infant death worldwide. Humans are the unique natural reservoir for this virus. It causes severe immunosuppression favouring secondary bacteri...
    • Natural killer cell mediated cytotoxicity, organism-specific biosystem (from KEGG)
      Natural killer cell mediated cytotoxicity, organism-specific biosystemNatural killer (NK) cells are lymphocytes of the innate immune system that are involved in early defenses against both allogeneic (nonself) cells and autologous cells undergoing various forms of stre...
    • Natural killer cell mediated cytotoxicity, conserved biosystem (from KEGG)
      Natural killer cell mediated cytotoxicity, conserved biosystemNatural killer (NK) cells are lymphocytes of the innate immune system that are involved in early defenses against both allogeneic (nonself) cells and autologous cells undergoing various forms of stre...

    Markers

    Homology

    Clone Names

    • FLJ18687, FLJ92177

    Gene Ontology Provided by GOA

    Function Evidence Code Pubs
    SH3/SH2 adaptor activity NAS
    Non-traceable Author Statement
    more info
    PubMed 
    Process Evidence Code Pubs
    cell-cell signaling IDA
    Inferred from Direct Assay
    more info
    PubMed 
    cellular defense response NAS
    Non-traceable Author Statement
    more info
    PubMed 
    humoral immune response IEA
    Inferred from Electronic Annotation
    more info
     
    positive regulation of natural killer cell mediated cytotoxicity IEA
    Inferred from Electronic Annotation
    more info
     
    Component Evidence Code Pubs
    cytoplasm IDA
    Inferred from Direct Assay
    more info
    PubMed 
    Preferred Names
    SH2 domain-containing protein 1A
    Names
    SH2 domain-containing protein 1A
    SLAM-associated protein
    Duncan disease SH2-protein
    T cell signal transduction molecule SAP
    T-cell signal transduction molecule SAP
    SLAM associated protein/SH2 domain protein 1A
    signaling lymphocyte activation molecule-associated protein
    signaling lymphocytic activation molecule-associated protein

    RefSeqs maintained independently of Annotated Genomes

    These reference sequences exist independently of genome builds. Explain

    These reference sequences are curated independently of the genome annotation cycle, so their versions may not match the RefSeq versions in the current genome build. Identify version mismatches by comparing the version of the RefSeq in this section to the one reported in Genomic regions, transcripts, and products above.

    Genomic

    1. NG_007464.1 RefSeqGene

      Range
      4983..31861
      Download
      GenBank, FASTA, Sequence Viewer (Graphics), LRG_106

    mRNA and Protein(s)

    1. NM_001114937.2NP_001108409.1  SH2 domain-containing protein 1A isoform 2

      Status: REVIEWED

      Description
      Transcript Variant: This variant (2) uses an alternate in-frame splice site in the 3' coding region, compared to variant 1, resulting in a shorter protein (isoform 2), compared to isoform 1.
      Source sequence(s)
      AA190204, AF100541, AL022718, AL023657, DA007747
      Consensus CDS
      CCDS48162.1
      UniProtKB/Swiss-Prot
      O60880
      Conserved Domains (1) summary
      cd10400
      Location:2104
      Blast Score: 535
      SH2_SAP1a; Src homology 2 (SH2) domain found in SLAM-associated protein (SAP) 1a
    2. NM_002351.4NP_002342.1  SH2 domain-containing protein 1A isoform 1

      Status: REVIEWED

      Description
      Transcript Variant: This variant (1) represents the longer transcript and encodes the longer isoform (1).
      Source sequence(s)
      AA190204, AL022718, AL023657
      Consensus CDS
      CCDS14608.1
      UniProtKB/Swiss-Prot
      O60880
      Related
      ENSP00000360181, OTTHUMP00000023976, ENST00000371139, OTTHUMT00000058174
      Conserved Domains (1) summary
      cd10400
      Location:2104
      Blast Score: 534
      SH2_SAP1a; Src homology 2 (SH2) domain found in SLAM-associated protein (SAP) 1a

    RefSeqs of Annotated Genomes: Homo sapiens Annotation Release 104

    The following sections contain reference sequences that belong to a specific genome build. Explain

    Reference GRCh37.p10 Primary Assembly

    Genomic

    1. NC_000023.10 Reference GRCh37.p10 Primary Assembly

      Range
      123480132..123507010
      Download
      GenBank, FASTA, Sequence Viewer (Graphics)

    Alternate HuRef

    Genomic

    1. AC_000155.1 Alternate HuRef

      Range
      112860780..112888184
      Download
      GenBank, FASTA, Sequence Viewer (Graphics)

    Alternate CHM1_1.0

    Genomic

    1. NC_018934.1 Alternate CHM1_1.0

      Range
      123434923..123461885
      Download
      GenBank, FASTA, Sequence Viewer (Graphics)

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