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    CDKN1C cyclin-dependent kinase inhibitor 1C (p57, Kip2) [ Homo sapiens ]

    Gene ID: 1028, updated on 13-May-2012

    Summary

    Official Symbol
    CDKN1Cprovided by HGNC
    Official Full Name
    cyclin-dependent kinase inhibitor 1C (p57, Kip2)provided by HGNC
    Primary source
    HGNC:1786
    See related
    Ensembl:ENSG00000129757; HPRD:02913; MIM:600856; Vega:OTTHUMG00000010040
    Gene type
    protein coding
    RefSeq status
    REVIEWED
    Organism
    Homo sapiens
    Lineage
    Eukaryota; Metazoa; Chordata; Craniata; Vertebrata; Euteleostomi; Mammalia; Eutheria; Euarchontoglires; Primates; Haplorrhini; Catarrhini; Hominidae; Homo
    Also known as
    BWS; WBS; p57; BWCR; KIP2
    Summary
    This gene is imprinted, with preferential expression of the maternal allele. The encoded protein is a tight-binding, strong inhibitor of several G1 cyclin/Cdk complexes and a negative regulator of cell proliferation. Mutations in this gene are implicated in sporadic cancers and Beckwith-Wiedemann syndorome, suggesting that this gene is a tumor suppressor candidate. Three transcript variants encoding two different isoforms have been found for this gene. [provided by RefSeq, Oct 2010]

    Genomic context

    Location :
    11p15.5
    Sequence :
    Chromosome: 11; NC_000011.9 (2904448..2906995, complement)
    See CDKN1C in Epigenomics, MapViewer

    Chromosome 11 - NC_000011.9Genomic Context describing neighboring genes Neighboring gene potassium voltage-gated channel, KQT-like subfamily, member 1 Neighboring gene KCNQ1 opposite strand/antisense transcript 1 (non-protein coding) Neighboring gene solute carrier family 22 (organic cation transporter), member 18 antisense Neighboring gene KCNQ1 downstream neighbor (non-protein coding) Neighboring gene solute carrier family 22, member 18 Neighboring gene pleckstrin homology-like domain, family A, member 2

    Genomic regions, transcripts, and products

    Bibliography

    GeneRIFs: Gene References Into Functions What's a GeneRIF?

    Phenotypes

    Beckwith-Wiedemann syndrome

    Summary from GeneReviews: Go to GeneReviews

    Disease Characteristics
    Beckwith-Wiedemann syndrome (BWS) is a growth disorder characterized by macrosomia, macroglossia, visceromegaly, embryonal tumors (e.g., Wilms tumor, hepatoblastoma, neuroblastoma, and rhabdomyosarcoma), omphalocele, neonatal hypoglycemia, ear creases/pits, adrenocortical cytomegaly, and renal abnormalities (e.g., medullary dysplasia, nephrocalcinosis, medullary sponge kidney, and nephromegaly). Early death may occur from complications of prematurity, hypoglycemia, cardiomyopathy, macroglossia, or tumors. However, the previously reported mortality of 20% is likely an overestimate given better recognition of the disorder along with enhanced treatment options. Macroglossia and macrosomia are generally present at birth but may have postnatal onset. Growth rate slows around age seven to eight years. Hemihyperplasia may affect segmental regions of the body or selected organs and tissues.
    Diagnosis Testing
    A provisional diagnosis of BWS based on clinical assessment may be confirmed by molecular/cytogenetic testing. Cytogenetically detectable abnormalities involving chromosome 11p15 are found in 1% or fewer of affected individuals. Clinically available molecular genetic testing can identify epigenetic and genomic alterations of chromosome 11p15 in individuals with BWS: (1) loss of methylation on the maternal chromosome at imprinting center 2 (IC2) in 50% of affected individuals; (2) paternal uniparental disomy for chromosome 11p15 in 20%; and (3) gain of methylation on the maternal chromosome at imprinting center 1 (IC1) in 5%. Methylation alterations that are associated with microdeletions or microduplications in this region are associated with high heritability. Sequence analysis of CDKN1C identifies mutations in approximately 40% of familial cases and 5%-10% of cases with no family history of BWS.
    Genetic Counseling
    Beckwith-Wiedemann syndrome is associated with abnormal regulation of gene transcription in the imprinted domain on chromosome 11p15.5. Most individuals with BWS are reported to have normal chromosome studies or karyotypes. Approximately 85% of individuals with BWS have no family history of BWS while approximately 15% have a family history consistent with autosomal dominant transmission of BWS. Children of subfertile parents conceived by assisted reproductive technology (ART) may have an increased risk for imprinting disorders, including BWS. Identification of the underlying genetic mechanism causing BWS permits better estimation of recurrence risk. Prenatal screening for pregnancies not previously known to be at increased risk for BWS by ultrasound examination and maternal serum alpha-fetoprotein assay may lead to the consideration of chromosome analysis and/or molecular genetic testing. Specific prenatal testing is possible by chromosome analysis for families with an inherited chromosome abnormality or by molecular genetic testing for families in which the molecular mechanism of BWS has been defined.
    References

    Common variants in the calcium-sensing receptor gene are associated with total serum calcium levels.

    Interactions

    Products Interactant Other Gene Complex Source Pubs Description
    NC_000011.8 PIAS2    BIND  PubMed Miz1 interacts with p-P57KIP2. 
    P49918 O96020 CCNE2    HPRD  PubMed  
    P49918 P11802 CDK4    HPRD  PubMed  
    P49918 P53667 LIMK1    HPRD  PubMed  
    P49918 P15172 MYOD1    HPRD  PubMed  
    P49918 P43354 NR4A2    HPRD  PubMed  
    P49918 P12004 PCNA    HPRD  PubMed  
    P49918 Q13309 SKP2    HPRD  PubMed  
    BioGRID:107462 BioGRID:107331 CCNA2    BioGRID  PubMed Affinity Capture-Western 
    BioGRID:107462 BioGRID:107067 CCND1    BioGRID  PubMed Reconstituted Complex 
    BioGRID:107462 BioGRID:114032 CUL1    BioGRID  PubMed Biochemical Activity 
    BioGRID:107462 BioGRID:198624 Cdk1    BioGRID  PubMed Affinity Capture-MS 
    BioGRID:107462 BioGRID:110172 LIMK1    BioGRID  PubMed Affinity Capture-Western; Two-hybrid 
    BioGRID:107462 BioGRID:110690 MYBL2    BioGRID  PubMed Affinity Capture-Western; Reconstituted Complex 
    BioGRID:107462 BioGRID:110737 MYOD1    BioGRID  PubMed Affinity Capture-Western; Reconstituted Complex 
    BioGRID:107462 BioGRID:111142 PCNA    BioGRID  PubMed Affinity Capture-Western; Reconstituted Complex; Two-hybrid 
    BioGRID:107462 BioGRID:111868 RBL1    BioGRID  PubMed Affinity Capture-Western 
    BioGRID:107462 BioGRID:112393 SKP2    BioGRID  PubMed Affinity Capture-Western 
    BioGRID:107462 BioGRID:113164 UBC    BioGRID  PubMed Affinity Capture-MS; Affinity Capture-Western 

    General gene information

    Markers

    Homology

    Pathways from BioSystems

    • Cell cycle, organism-specific biosystem (from KEGG)
      Cell cycle, organism-specific biosystemMitotic cell cycle progression is accomplished through a reproducible sequence of events, DNA replication (S phase) and mitosis (M phase) separated temporally by gaps known as G1 and G2 phases. Cycli...
    • Cell cycle, conserved biosystem (from KEGG)
      Cell cycle, conserved biosystemMitotic cell cycle progression is accomplished through a reproducible sequence of events, DNA replication (S phase) and mitosis (M phase) separated temporally by gaps known as G1 and G2 phases. Cycli...
    • Endochondral Ossification, organism-specific biosystem (from WikiPathways)
      Endochondral Ossification, organism-specific biosystem
      Endochondral Ossification
    • G1 to S cell cycle control, organism-specific biosystem (from WikiPathways)
      G1 to S cell cycle control, organism-specific biosystemIn the G1 phase there are two types of DNA damage responses, the p53-dependent and the p53-independent pathways. The p53-dependent responses inhibit CDKs through the up-regulation of genes encoding C...

    Gene Ontology Provided by GOA

    Function Evidence Code Pubs
    cyclin-dependent protein kinase inhibitor activity IEA
    Inferred from Electronic Annotation
    more info
     
    protein binding IPI
    Inferred from Physical Interaction
    more info
    PubMed 
    protein kinase inhibitor activity IEA
    Inferred from Electronic Annotation
    more info
     
    Component Evidence Code Pubs
    cytoplasm IDA
    Inferred from Direct Assay
    more info
    PubMed 
    NOT nucleolus IDA
    Inferred from Direct Assay
    more info
    PubMed 
    nucleus IDA
    Inferred from Direct Assay
    more info
    PubMed 

    General protein information

    Preferred Names
    cyclin-dependent kinase inhibitor 1C
    Names
    cyclin-dependent kinase inhibitor 1C
    p57Kip2
    cyclin-dependent kinase inhibitor p57

    NCBI Reference Sequences (RefSeq)

    RefSeqs maintained independently of Annotated Genomes

    These reference sequences exist independently of genome builds. Explain

    These reference sequences are curated independently of the genome annotation cycle, so their versions may not match the RefSeq versions in the current genome build. Identify version mismatches by comparing the version of the RefSeq in this section to the one reported in Genomic regions, transcripts, and products above.

    Genomic

    1. NG_008022.1 RefSeqGene

      Range
      5001..7548
      Download
      GenBank, FASTA, Sequence Viewer (Graphics)

    mRNA and Protein(s)

    1. NM_000076.2NP_000067.1  cyclin-dependent kinase inhibitor 1C isoform a

      Status: REVIEWED

      Description
      Transcript Variant: This variant (1) represents the longest transcript and encodes the longer isoform (a).
      Source sequence(s)
      AC013791, BC039188, CD676249, U22398
      Consensus CDS
      CCDS7738.1
      UniProtKB/Swiss-Prot
      P49918
      Related
      ENSP00000413720, OTTHUMP00000011736, ENST00000414822, OTTHUMT00000027774
      Conserved Domains (1) summary
      pfam02234
      Location:3183
      Blast Score: 173
      CDI; Cyclin-dependent kinase inhibitor
    2. NM_001122630.1NP_001116102.1  cyclin-dependent kinase inhibitor 1C isoform b

      Status: REVIEWED

      Description
      Transcript Variant: This variant (2) differs in the 5' UTR and coding sequence compared to variant 1. The resulting isoform (b) is shorter at the N-terminus compared to isoform a. Variants 2 and 3 both encode isoform b.
      Source sequence(s)
      AC013791, BC039188, CD676249, U22398
      Consensus CDS
      CCDS44519.1
      UniProtKB/Swiss-Prot
      P49918
      Related
      ENSP00000411257, OTTHUMP00000011738, ENST00000440480, OTTHUMT00000027776
      Conserved Domains (1) summary
      pfam02234
      Location:2072
      Blast Score: 170
      CDI; Cyclin-dependent kinase inhibitor
    3. NM_001122631.1NP_001116103.1  cyclin-dependent kinase inhibitor 1C isoform b

      Status: REVIEWED

      Description
      Transcript Variant: This variant (3) differs in the 5' UTR and coding sequence and uses an alternate splice junction in the 3' UTR compared to variant 1. The resulting isoform (b) is shorter at the N-terminus compared to isoform a. Variants 2 and 3 both encode isoform b.
      Source sequence(s)
      AC013791, BC039188, BM673714, CD676249, U22398
      Consensus CDS
      CCDS44519.1
      UniProtKB/Swiss-Prot
      P49918
      Related
      ENSP00000321019, ENST00000313407
      Conserved Domains (1) summary
      pfam02234
      Location:2072
      Blast Score: 170
      CDI; Cyclin-dependent kinase inhibitor

    RefSeqs of Annotated Genomes: Build 37.3

    The following sections contain reference sequences that belong to a specific genome build. Explain

    Reference GRCh37.p5 Primary Assembly

    Genomic

    1. NC_000011.9 Reference GRCh37.p5 Primary Assembly

      Range
      2904448..2906995, complement
      Download
      GenBank, FASTA, Sequence Viewer (Graphics)

    Alternate HuRef

    Genomic

    1. AC_000143.1 Alternate HuRef

      Range
      2692706..2695485, complement
      Download
      GenBank, FASTA, Sequence Viewer (Graphics)

    Related Sequences

    Nucleotide Protein
    Heading Accession and Version
    genomic AC013791.9 (56638..58748) None
    genomic CH471158.1 EAX02524.1
      EAX02525.1
    genomic D64137.1 BAA11014.1
      BAA11015.1
    genomic U48869.1 AAB05896.1
    mRNA BC039188.1 None
    mRNA BC067842.1 AAH67842.1
    mRNA BM673714.1 None
    mRNA CD676249.1 None
    mRNA U22398.1 AAA85095.1
    Protein Accession Links
    GenPept Link UniProtKB Link
    P49918.1 GenPept UniProtKB/Swiss-Prot:P49918

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